1. Division of Biological Science, Nara Institute of Science and Technology, Ikoma, Nara, Japan
2. Department of Cellular Physiology, Niigata University Graduate School of Medical and Dental Sciences, Niigata, Japan
3. Department of Life Sciences, Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan
4. National Institute for Basic Biology, Okazaki, Aichi, Japan
5. Graduate University for Advanced Studies (SOKENDAI), Hayama, Kanagawa, Japan
6. Department of Microbiology and Molecular Genetics, University of California, Davis, California, USA