1. Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, 610041, P.R. China
2. Graduate University of the Chinese Academy of Sciences, Beijing, 100049, P.R. China
3. Key Laboratory of Ecological Restoration, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, 610041, P.R. China
4. Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chinese Academy of Sciences, Chengdu, 610041, P.R. China