1. Department of Ocean Science and Hong Kong Branch of Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Hong Kong University of Science and Technology, Hong Kong, Hong Kong
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou, China
3. Department of Biological Sciences, Université de Montréal, Montréal, Canada
4. Center of Deep Sea Research Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China