1. College of Life Science, Neijiang Normal University, Neijiang, China;
2. Key Laboratory of Freshwater Fish Reproduction and Development, Ministry of Education Laboratory of Aquatic Science of Chongqing, School of Life Sciences, Southwest University, Chongqing, China;
3. Key Laboratory of Sichuan Province for Fishes Conservation and Utilization in the Upper Reaches of the Yangtze River, Neijiang Normal University, Neijiang, China;
4. BGI Education Center, University of Chinese Academy of Sciences, Shenzhen, China;