1. Qinghai Provincial Key Laboratory of High value Utilization of Characteristic Economic Plants, The College of Ecological Environment and Resources, Qinghai Minzu University, Xining, Qinghai, P. R. China
2. Key Laboratory of Cold Regions Restoration Ecology, Qinghai Province, and Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, Qinghai, P. R. China