1. Laboratory of Insect Ecology, Graduate School of Agriculture, Kyoto University, Kyoto, Japan
2. Department of Evolutionary and Environmental Biology, Institute of Evolution, University of Haifa, Haifa, Israel
3. Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA, USA
4. Department of General Systems Studies, Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan