1. National Engineering Research Center for Marine Aquaculture, Zhejiang Ocean University, Zhoushan, Zhejiang, China
2. CAS Key Laboratory of Marine Ecology and Environmental Sciences (KLMEES), Institute of Oceanology, Chinese Academy of Sciences (IOCAS), Qingdao, Shandong, China
3. Marine Ecology and Environmental Science Laboratory, Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao, Shandong, China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, Shandong, China