1. Department of Biology, Faculty of Sciences, University of Isfahan, Isfahan, Iran
2. Natural Resources Research Department, Isfahan Agricultural and Natural Resources Research and Education Centre, Agricultural Research, Education and Extension Organisation (AREEO), Isfahan, Iran
3. Botany Research Division, Research Institute of Forests and Rangelands, Agricultural Research, Education and Extension Organisation (AREEO), Tehran, Iran
4. Leibniz Institute of Plant Genetics and Crop Plant Research (IPK), Gatersleben, Germany
5. German Centre of Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Leipzig, Germany