1. NIHR Nottingham Biomedical Research Centre, University of Nottingham, Nottingham, UK
2. Nottingham Digestive Diseases Centre, School of Medicine, University of Nottingham, Nottingham, UK
3. Biological Sciences, Clemson University, Clemson, SC, USA
4. Division of Digestive Diseases, Department of Metabolism, Digestion and Reproduction, Faculty of Medicine, Imperial College London, London, UK
5. Departments of Gastroenterology and Hepatology, St Mary’s Hospital, Imperial College Healthcare NHS Trust, London, UK
6. Department of Infection Biology, London School of Hygiene and Tropical Medicine, London, UK
7. Department of Gastroenterology, Zeidler Ledcor Centre, University of Alberta, Edmonton, Alberta, Canada
8. School of Pharmacy, Biodiscovery Institute, University of Nottingham, University Park, Nottingham, UK