1. Department of Medicine, University of Alberta, Edmonton, AB T6G 2R3, Canada
2. Faculty of Veterinary Medicine, University of Calgary, Calgary, AB T2N 4Z6, Canada
3. National Institute for Health Research, Nottingham Biomedical Research Centre, Nottingham Digestive Diseases Centre, School of Medicine, University of Nottingham, Nottingham NG7 2UH, UK
4. Faculty of Medicine and Dentistry, University of Alberta, Edmonton, AB T6G 2R3, Canada
5. Public Health Laboratory, Alberta Precision Laboratories, Edmonton, AB T6G 2R3, Canada