1. State Key Laboratory for Biocontrol, Southern Laboratory of Ocean Science and Engineering, Zhuhai, Guangdong, PR China
2. School of Marine Sciences, Sun Yat-sen University, Zhuhai, Guangzhou, PR China
3. Guangdong Province Key Laboratory for Aquatic Economic Animals, and Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, Sun Yat-sen University, Guangzhou, PR China