1. CAS Key Laboratory of Special Pathogens and Biosafety, Center for Emerging Infectious Diseases, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, China
2. College of life sciences and health, Wuhan university of science and technology, Wuhan, China
3. College of Veterinary Medicine, Huazhong Agricultural University, Wuhan, China
4. State Key Laboratory of Virology, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, P.R. China
5. Beijing Institute of Infectious Diseases,Military Hospital of China, Beijing, P.R. China