1. Laboratory for Parkinson's Disease, Tokyo Metropolitan Institute of Medical Science , Setagaya-ku, Japan
2. PCND Neuroscience Research Institute , Poway, CA, USA
3. Department of Physiology, Nippon Medical School , Tokyo, Japan
4. Division of Neurosciences, National Institute on Aging, National Institutes of Health , Bethesda, MD, USA
5. Institute of Agrobiological Sciences, National Agriculture and Food Research Organization , Tsukuba, Japan