1. State Key Laboratory of Forest and Soil Ecology, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, China
2. Key Laboratory for Management of Non-commercial Forests, Shenyang 110016, China
3. Qingyuan Forest CERN, Chinese Academy of Sciences, Shenyang 110016, China
4. Graduate University of Chinese Academy of Sciences, Beijing 100049, China