1. a Department of Environmental Biology and Fisheries Science , National Taiwan Ocean University , Keelung , 20224 , Taiwan
2. b Centre of Excellence for Marine Bioenvironment and Biotechnology , National Taiwan Ocean University , Keelung , 20224 , Taiwan
3. c National Research Institute of Far Seas Fisheries, Fisheries Research Agency , Shizuoka City , Shizuoka , 424-8633 , Japan
4. d Coastal and Offshore Resources Research Centre, Fisheries Research Institute, Council of Agriculture, Executive , Kaohsiung , 50672 , Taiwan