1. School of Science, the University of New South Wales, Canberra, Australia
2. Institute for Applied Ecology, University of Canberra, Canberra, Australia
3. Lund Vision Group, Department of Biology, Lund University, Lund, Sweden
4. Research School of Biology, Australian National University, Canberra, Australia
5. Division of Information, Technology and Development, University of South Australia, Adelaide, Australia