1. Plant and Invertebrate Ecology Department, Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, UK.
2. Department of Biology, University of York, Heslington, York YO10 5YW, UK.
3. Met Office, FitzRoy Road, Exeter EX1 3PB, UK.
4. School of Geography, University of Exeter, Exeter, Devon EX4 8RJ, UK.
5. Natural Resources Institute, University of Greenwich, Chatham, Kent ME4 4TB, UK.