1. RIKEN BioResource Research Center, Tsukuba, Ibaraki, Japan
2. Institute of Laboratory Animals, Kyoto University Graduate School of Medicine, Kyoto, Japan
3. Department of Anatomy and Embryology, University of Tsukuba, Ibaraki, Japan
4. Department of Animal Genetic Resources, National Gene Bank, Giza, Egypt
5. Graduate School of Life and Environmental Sciences, University of Tsukuba, Ibaraki, Japan
6. RIKEN Cluster for Pioneering Research, Saitama, Japan