1. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, U.S.A.;
2. pFriem Family Brewers, Hood River, OR, U.S.A.;
3. Department of Food Science and Technology, Oregon State University, Corvallis, OR, U.S.A.;
4. Department of Plant Science, Pennsylvania State University, University Park, PA, U.S.A.;
5. Deschutes Brewery, Bend, OR, U.S.A.;
6. Hartwick College Center for Craft Food and Beverage, Oneonta, NY, U.S.A.