1. Creative Research Institution, Hokkaido University , Kita-ku, Sapporo, 001-0021, Japan
2. Biotechnology Research Center, University of Tokyo , Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan
3. Graduate School of Agricultural and Life Sciences, The University of Tokyo , Tokyo, 113-8657, Japan
4. Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency , Saitama 332-0012, Japan