1. U.S. Geological Survey, Western Fisheries Research Center, 5501 A Cook-Underwood Road, Cook, Washington 98605, USA
2. School of the Environment and School of Biological Sciences, Washington State University, 14204 Northeast Salmon Creek Avenue, Vancouver, Washington 98686, USA
3. U.S. Geological Survey, Western Fisheries Research Center, 6505 Northeast 65th Street, Seattle, Washington 98115, USA