1. School of Earth Sciences, China University of Geosciences (Wuhan), No. 388 Lumo Road, Wuhan 430074, China,
2. Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142 Xizhimenwai Street, Beijing 100044, China, ,
3. Department of Biological Sciences, George Washington University, 2029 G Street, NW, Washington, D.C., 20052, U.S.A.,