1. Laboratory of Aromatic and Medicinal Plants, Center of Biotechnology, Technopark of Borj Cedria, Hammam-Lif, Tunisia
2. Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan
3. Alliance for Research on the Mediterranean and North Africa (ARENA), University of Tsukuba, Tsukuba, Japan
4. Faculty of Life and Environmental Sciences, University of Tsukuba, Tennodai, Tsukuba, Ibaraki, Japan