1. a Department of Biological Sciences , University of Warwick , Coventry , CV4 7AL , UK
2. b Fachbereich Biologie, Humboldt-Universität zu Berlin , Invalidenstrasse 43, D-10115 , Berlin , Germany
3. c Station Biologique de Roscoff , CNRS et Université Paris 6 , BP 74, F-29682 , Roscoff Cedex , France