1. Division of Histology and Embryology; Key Laboratory for Regenerative Medicine of the Ministry of Education; Medical College; Jinan University; Guangzhou, China
2. Department of Pathophysiology; Institute of Brain Research; Medical College; Jinan University; Guangzhou, China
3. The authors contributed equally to this work.
4. Division of Cell and Developmental Biology; University of Dundee; Dundee, UK
5. Key Laboratory of Medical Cell Biology, China Medical University, Shenyang, China
6. Institute of Vascular Biological Sciences; Guangdong Pharmaceutical University; Guangzhou, China