1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China
2. Chinese Academy of Sciences, Center for Excellence in Life and Palaeoenvironment, Beijing, China
3. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, China
4. Department of Anatomy, Histology and Embryology, Capital Medical University, Beijing, China
5. Department of Paleoanthropology, Yunnan Institute of Cultural Relics and Archaeology, Kunming, Yunnan, China