1. Department of Systematic Botany, Evolutionary Biology Centre, Uppsala University Norbyvägen 18 D, SE-752 36 Uppsala, Sweden; E-mail: sylvain.razafimandimbison@ebc.uu.se (S.G.R.)
2. The Bergius Foundation at the Royal Swedish Academy of Sciences P.O. Box 50017, SE-104 05 Stockholm, Sweden
3. University of Missouri–St. Louis 8001 Natural Bridge Road, St. Louis, Missouri 63121, USA