1. Kasuya Research Forest, Kyushu University , Sasaguri, Japan
2. Miyazaki University Forests, University of Miyazaki , Tano, Japan
3. Department of Environmental Sciences and Technology, Kagoshima University , Kagoshima, Japan
4. Graduate School of Agriculture, Ehime University , Matusyama, Japan
5. Graduate School of Agricultural Science, Kobe University , Kobe, Japan