1. a Key Laboratory of Zoological Systematics and Evolution , Institute of Zoology, Chinese Academy of Sciences , Box 92, No. 1, Beichen West Road, Chaoyang District, Beijing , 100101 , PR China
2. b Institut für Spezielle Zoologie und Evolutionsbiologie , FSU Jena , Germany
3. c College of Life Sciences , Capital Normal University , Beijing , 100048 , PR China