1. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China. ; dyhuang@nigpas.ac.cn; cycai@nigpas.a
2. School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS8 1TQ, UK.
3. Department of Integrative Biology, Oregon State Arthropod Collection, Oregon State University, Corvallis, OR 97331, USA. ; david.maddison@oregonstate.edu
4. Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.