1. Department of Biological Sciences, The George Washington University, Washington, DC 20052, USA;
2. Department of Anatomical Sciences, Stony Brook University, Stony Brook, NY 11794-8081, USA;
3. Department of Earth Sciences, University College London, London WC1E 6BS, UK;
4. Department of Earth Sciences, Natural History Museum, London SW7 5BD, UK;
5. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, People’s Republic of China