1. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, South China Botanical Garden, Chinese Academy of Sciences, No.723, Xingke Road, Tianhe District, Guangzhou 510650, China;
2. Center of Plant Ecology, Core Botanical Gardens, Chinese Academy of Sciences, No.723, Xingke Road, Tianhe District, Guangzhou 510650, China;
3. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences, No.723, Xingke Road, Tianhe District, Guangzhou 510650, China;
4. Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555, boulevard de l’Université, Chicoutimi G7H 2B1, Canada