1. Graduate School of Integrated Sciences for Life, Hiroshima University, Higashi-Hiroshima, Japan
2. Faculty of Agriculture and Marine Science, Kochi University, Kochi, Japan
3. Ion Beam Breeding Team Beam Mutagenesis Group, RIKEN Nishina Center for Accelerator-Based Science, Wako, Japan
4. Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan
5. Department of Plant Breeding, Plant Breeding, Wageningen University and Research, Wageningen, The Netherlands