1. Department of Botany, 3190 Maile Way, University of Hawaiʻi, Honolulu, HI 96822, USA
2. Marine Biology Graduate Program, University of Hawaiʻi, Honolulu, HI 96822, USA
3. Institute of Marine Biology, National Taiwan Ocean University, Keelung 20224, Taiwan, R.O.C.
4. Department of Biological Sciences, 3209 N. Maryland Avenue, University of Wisconsin Milwaukee, Milwaukee, WI 53212, USA
5. Department of Biology, College of Charleston, 66 George St, Charleston, SC 29424, USA
6. NOAA, Papahānaumokuākea Marine National Monument, 1845 Wasp Boulevard, Building 176, Honolulu, HI 96818, USA