1. Laboratory of Applied Phycology, Graduate School of Tokyo University of Marine Science and Technology, 4-5-7 Konan, Minato-ku, Tokyo, Japan
2. National Research Institute of Fisheries Science, Fukuura 2-12-4, Kanazawa-ku, Yokohama, Kanagawa, Japan
3. AXIOHELIX Co. Ltd, Nihonbashi, Chuou-ku, Tokyo, 103‐0015 Japan