1. National Institute of Water and Atmospheric Research Ltd, Private Bag 14-901, Wellington 6241, New Zealand
2. Institute of Marine Biology, National Taiwan Ocean University, Keelung 20224, Taiwan, Republic of China
3. Herbarium and Biology Department, University of North Carolina – Chapel Hill, Coker Hall, CB 3280, Chapel Hill, North Carolina 27599-3280, USA
4. School of Biological Sciences, Victoria University of Wellington, PO Box 600, Wellington, New Zealand