1. Kobe University Research Center for Inland Seas, Rokkodai, Nadaku, Kobe 657-8501, Japan
2. Graduate School of Science, Kobe University, Rokkodai, Nadaku, Kobe 657-8501, Japan; cUniversity Herbarium, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, CA 94720, USA
3. Bezhin Rosko, 40 rue des pêcheurs, 29250 Santec, France