1. Kobe University Research Center for Inland Seas, Rokkodai, Nadaku, Kobe 657-8501, Japan
2. Graduate School of Science, Kobe University, Rokkodai, Kobe 657-8501, Japan
3. School of Marine Biosciences, Kitasato University, Kitazato, Minamiku, Sagamihara 252-0373, Japan
4. Science, Natural History Museum, Cromwell Road, London SW7 5BD, UK
5. University Herbarium, University of California, Berkeley, CA 94720, USA
6. National Institute of Water & Atmospheric Research, Wellington 6241, New Zealand
7. School of Biological Sciences, University of Auckland, Auckland 1142, New Zealand