1. University of Girona, Faculty of Sciences, Department of Environmental Sciences, M. Aurèlia Capmany 69, 17003 Girona, Spain
2. Phycology Research Group and Centre for Molecular Phylogenetics and Evolution, Ghent University, Ghent, Belgium
3. Western Australian Herbarium, Biodiversity and Conservation Science, Department of Biodiversity, WA, 6983, Australia
4. Institute of Marine Biology, National Taiwan Ocean University, Keelung 20224, Taiwan, R.O.C.