1. Faculty of Sciences, Department of Environmental Sciences, University of Girona, M. Aurèlia Capmany 69, Girona 17003, Spain
2. Western Australian Herbarium, Biodiversity and Conservation Science, Department of Biodiversity, Conservation and Attractions, Locked Bag 104, Bentley Delivery Centre, WA 6983, Australia
3. Institute of Marine Biology, National Taiwan Ocean University, Keelung 20224, Taiwan, R.O.C