1. Kirstenbosch Research Centre, South African National Biodiversity Institute, Cape Town, South Africa
2. Department of Biological Sciences, University of Cape Town, Rondebosch, South Africa
3. Statistics in Ecology, Environment and Conservation, Department of Statistical Sciences, University of Cape Town, Rondebosch, South Africa
4. Department of Ancient Studies, Stellenbosch University, Stellenbosch, South Africa
5. CIRAD, UMR PVBMT, La Réunion, France