1. Research Laboratory of Environmental Sciences and Technologies (LR16ES09), Higher Institute of Environmental Sciences and Technology, University of Carthage, Hammam-Lif, Tunisia
2. Laboratory of Microorganisms and Biomolecules of the Center of Biotechnology of Sfax-Tunisia, Sfax, Tunisia
3. Zoology Department, College of Science, King Saud University, Saudi Arabia, Riyadh, Saudi Arabia