1. Department of Applied Biological Science, Interdisciplinary Graduate School of Agriculture and Engineering, University of Miyazaki, Miyazaki, Japan
2. Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, Yaeyama-gun, Okinawa 907-1541, Japan
3. Faculty of Agriculture, Department of Marine Biology and Environmental Science, University of Miyazaki, Miyazaki, Japan