1. International Center for Island Studies, Kagoshima University, Naze-Minatomachi 15-1-6F, Amami, Kagoshima 894-0026, Japan
2. Molecular Invertebrate Systematics and Ecology Laboratory, Graduate School of Engineering and Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan