1. Misaki Marine Biological Station, The University of Tokyo, Koajiro 1024, Misaki, Miura, Kanagawa 238-0225, Japan
2. Graduate School of Science, Hokkaido University, N10E8 Kita-ku, Sapporo, Hokkaido 060-0180, Japan
3. Faculty of Science, Hokkaido University, N10E8 Kita-ku, Sapporo, Hokkaido 060-0180, Japan