1. Faculty of Science, Hokkaido University, Kita-ku N10E8, Sapporo, Hokkaido 060-0810, Japan
2. Graduate School of Science, Hokkaido University, Kita-ku N10E8, Sapporo, Hokkaido 060-0810, Japan
3. Misaki Marine Biological Station, Graduate School of Science, The University of Tokyo, 1024 Koajiro, Misaki, Miura, Kanagawa 238-0225, Japan