1. Barel, C. D. N., F. Witte & M. J. P. van Oijen, 1976. The shape of the skeletal elements in the head of a generalized Haplochromis species: H. elegans Trewavas 1933 (Pisces, Cichlidae). Netherlands Journal of Zoology 26: 163–265.
2. Barel, C. D. N., M. J. P. van Oijen, F. Witte & E. L. M. Witte-Maas, 1977. An introduction to the taxonomy and morphology of the haplochromine cichlids from Lake Victoria. Netherlands Journal of Zoology 27: 333–389.
3. Boulenger, G. A., 1906. Descriptions of new fishes discovered by Mr. E. Degen in Lake Victoria. Annals and Magazine of Natural History 17: 433–452.
4. Boulenger, G. A., 1911. On a third collection of fishes made by Dr. E. Bayon in Uganda, 1909–1910. (Collezioni zoologiche fatte nell-Uganda dal Dott. E. Bayon. XII.). Annali del Museo Civico di Storia Naturale de Genova 5: 64–78.
5. Boulenger, G. A., 1914. Fische. In Wissenschaftlichen ergebnisse der Deutschen Zentral-Afrika Expedition, 1907–1908, Zoologie,Vol. 3: 354.