1. Centre of Research in Biodiversity, Ecology, Evolution and Conservation (CRBEC), DRC
2. Département de Biologie-Chimie, Unité d’Enseignement et de Recherche en Hydrobiologie Appliquée, Institut Supérieur Pédagogique (ISP) of Bukavu, Bukavu, 854, DRC
3. 3Vertebrates section, Ichthyology, Royal Museum for Central Africa (RMCA), Leuvensesteenweg 13, Tervuren, 3080, Belgium
4. Biology Department, Fish Diversity and Conservation, KU Leuven, Charles Deberiotstraat 32, Leuven, B-3000, Belgium
5. Zoological Institute, University of Basel, Vesalgasse 1, Basel, 4051, Switzerland
6. Faculty of Biology, Division of Evolutionary Biology, LMU Munich, Großhaderner Straße 2, Planegg-Martinsried, 82152, Germany