1. grid.263488.3 0000 0001 0472 9649 Shenzhen Key Laboratory of Microbial Genetic Engineering, College of Life Sciences and Oceanography Shenzhen University 518060 Shenzhen People’s Republic of China
2. grid.412067.6 0000 0004 1760 1291 Engineering Research Center of Agricultural Microbiology Technology, Ministry of Education Heilongjiang University 150500 Harbin People’s Republic of China
3. grid.495261.d 0000 0004 1797 8750 College of Culture and Media Hezhou University 542800 Hezhou People’s Republic of China
4. grid.412067.6 0000 0004 1760 1291 Key Laboratory of Molecular Biology, College of Heilongjiang Province, School of Life Sciences Heilongjiang University 150080 Harbin People’s Republic of China