1. School of Life Sciences, University of KwaZulu-Natal, Durban, South Africa
2. Current affiliation: Geological Sciences, School of Agricultural, Earth and Environmental Sciences, University of KwaZulu-Natal, Durban, South Africa
3. Department of Biodiversity and Conservation Biology, University of the Western Cape, Bellville, South Africa
4. Oceanographic Research Institute, Durban, South Africa
5. Branch: Oceans and Coasts, Department of Environmental Affairs, Cape Town, South Africa
6. Marine Research Institute, Department of Biological Sciences, University of Cape Town, Cape Town, South Africa
7. Medgar Evers College, City University of New York, New York, USA